Exchange server 2007 stop receiving emails

i have a strange issue with our exchange 2007 server, and i hope any one can help me, each day i have one of our exchange servers can't receive emails, and all emails sent to users in this servers are queued in the sending email server, and i have to manually
restart the Microsoft exchange transport service in this server so that it's able to receive emails, i found an error in the server that always repeated with different parameters
the error is
Error ID: 4999
Error Source: MSExchange Common
Watson report about to be sent to dw20.exe for process id: 3508, with parameters: E12, c-RTL-AMD64, 08.02.0254.000, edgetransport, M.E.Transport, M.E.P.S.SmtpInServer.HandleConnection, System.NullReferenceException, 69f8, 08.02.0254.000.  ErrorReportingEnabled:
False
i receive the same error each time the server stop responds but the parameters is changed each time (Process ID number always changed) like below
Watson report about to be sent to dw20.exe for process id: 3644, with parameters: E12, c-RTL-AMD64, 08.02.0254.000, edgetransport, M.E.Transport, M.E.P.S.SmtpInServer.HandleConnection, System.NullReferenceException, 69f8, 08.02.0254.000.  ErrorReportingEnabled:
False
i hope any one can help me solving this problem
the server is windows server 2008
Exchange 2007 SP2 Server Rollup 5
McAfee Enterprise Antivirus is in the box
Backup Exec Agent is in the box Also
a lot of transport rules applied to our exchange organization
Mahmoud Sabry IT System Engineer

Hi
As per the information and details provided by you, Exchange server 2007 stop receiving emails, then please follow these steps: -
If you do have a 60 GB partition on the C: drive, one of the first signs of trouble is when the free space on C: drops below 2GB. Exchange 2007 stops receiving external
emails. This is caused by a condition known as Back Pressure.
Please check the following: -
Check the nslook up for correct MX record.
Run the exbpa and llok for any recent changes.
Check for the device in perimeter network which accepts connections on port 25 FW or any other anti spam device).
Try to run a telnet session from the firwall and then out side and drop email on the server.
I hope this information will be helpful for you.
Thanks and regards
Shweta@G 

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