HT3918 Can I leave the installed 4gb (2 X 2gb) of ram in my iMac and add 8 gb (2 X 4gb) of additional ram to the available (empty) slots?

Can I leave the installed 4gb (2 X 2gb) of ram in my iMac and add 8 gb (2 X 4gb) of additional ram to the available (empty) slots? I will ultimately swap the remaining 2 X 2gb out for 2 x 4gb.  If so, does this affect how the ram is used/accessed (i.e. will the original ram be limiting because it is in the installed location rather than in the other previously open slots?)  Thanks. 

That's exactly the RAM arrangement I've been running with for the last couple of years with no issues.
2x2gb on one shelf and 2x4gb on the second.

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