MacBook Air can't connect to iMac network, but other computers can

This one has been driving me absolutely batty.
I have 4 computers on our network – a mini, iMac, Macbook, and Macbook Air. Screen and file sharing works great except everywhere except between the iMac and Air, which can't do anything with each other. They show up in each other's "Devices" list in the Finder, but trying to connect over any port, pinging, etc. doesn't work.
I've tried turning off the Firewall, changing IP addresses and dns settings, rebooting, turning off ipv6, creating new user profiles on both machines, and just about anything else I can think of. What am I missing
Thank you kindly for any help you can provide!

That's been one of the more difficult parts – no connections work over any protocols/ports that I've tried, whether via browsing or Connect to Server..., though something is working (UDP?) to allow both computers to see each other in the devices list in the Finder.
It happened just now. I was working on the Air and had mounted the iMac. It worked fine for about 15 minutes and then I suddenly got the pinwheel of death when trying to access the mounted iMac volume. Hanging, hanging, hanging, so I relaunched the Finder. Now I can't access the iMac at all (except, again, to see it in the devices list).
Here are seemingly related messages in the console (note that there is 1.2 TB unused on iMac and 11 GB unused on Air). It's showing some kind of file system error. Guess I'll run fsck and see what happens.
Thanks again for your help!
3/2/11 9:44 PM KernelEventAgent[37] tid 00000000 received event(s) VQ_LOWDISK, VQ_VERYLOWDISK (516)
3/2/11 9:45 PM hdiejectd[5532] quitCheck: calling exit(0)
3/2/11 9:58 PM quicklookd[6053] [QL] 'Creating thumbnail' timed out for '<QLThumbnailRequest /Volumes/jed/Documents/lost/food.eps>' (Start date: 2011-03-02 21:57:41 -0500)
3/2/11 9:58 PM KernelEventAgent[37] tid 00000000 received event(s) VQ_NOTRESP (1)
3/2/11 9:58 PM KernelEventAgent[37] tid 00000000 type 'afpfs', mounted on '/Volumes/jed', from 'afp_0X76lf1xMNqb4yHcGc0cKW9p-1.2d000006', not responding
3/2/11 9:58 PM KernelEventAgent[37] tid 00000000 found 1 filesystem(s) with problem(s)
3/2/11 9:58 PM kernel ASP_TCP Disconnect: triggering reconnect by bumping reconnTrigger from curr value 0 on so 0x70cab28
3/2/11 9:58 PM kernel AFP_VFS afpfs_DoReconnect started /Volumes/jed prevTrigger 0 currTrigger 1
3/2/11 9:58 PM kernel AFP_VFS afpfs_DoReconnect: doing reconnect on /Volumes/jed
3/2/11 9:58 PM kernel AFP_VFS afpfs_DoReconnect: posting to KEA EINPROGRESS for /Volumes/jed
3/2/11 9:58 PM kernel AFP_VFS afpfs_DoReconnect: Max reconnect time: 30 secs, Connect timeout: 15 secs for /Volumes/jed
3/2/11 9:58 PM kernel AFP_VFS afpfs_DoReconnect: connect to the server /Volumes/jed
3/2/11 9:59 PM kernel AFP_VFS afpfs_DoReconnect: connect on /Volumes/jed failed 60.
3/2/11 9:59 PM kernel AFP_VFS afpfs_DoReconnect: connect to the server /Volumes/jed
3/2/11 9:59 PM kernel AFP_VFS afpfs_DoReconnect: connect on /Volumes/jed failed 60.
3/2/11 9:59 PM kernel ]
3/2/11 9:59 PM kernel AFP_VFS afpfs_DoReconnect: posting to KEA to unmount /Volumes/jed
3/2/11 9:59 PM kernel ASP_TCP CancelOneRequest: cancelling slot 0 error 89 reqID 6337 flags 0x29 afpCmd 0x3C so 0x70cab28
3/2/11 9:59 PM kernel ASP_TCP CancelOneRequest: cancelling slot 1 error 89 reqID 6338 flags 0x29 afpCmd 0x22 so 0x70cab28
3/2/11 9:59 PM kernel ASP_TCP Disconnect: triggering reconnect by bumping reconnTrigger from curr value 1 on so 0x70cab28
3/2/11 9:59 PM kernel AFP_VFS afpfs_DoReconnect started /Volumes/jed prevTrigger 1 currTrigger 2
3/2/11 9:59 PM kernel AFP_VFS afpfs_DoReconnect: already in unmount /Volumes/jed
3/2/11 9:59 PM kernel
3/2/11 9:59 PM KernelEventAgent[37] tid 00000000 received event(s) VQ_DEAD (32)
3/2/11 9:59 PM KernelEventAgent[37] tid 00000000 type 'afpfs', mounted on '/Volumes/jed', from 'afp_0X76lf1xMNqb4yHcGc0cKW9p-1.2d000006', dead
3/2/11 9:59 PM KernelEventAgent[37] tid 00000000 found 1 filesystem(s) with problem(s)

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