What is the box in mail app between the move icon and reply icon?

Hello,
Just started using mail in the new ipad and see an icon in the inbox on the left handside in between the move icon and the reply icon. What is it and what does it do to a file? I clicked an old email on it and it seemed to have stored it or did it delete it. Thanks for your help in this matter.

Do you mean the icons towards the top right-hand side of the Mail app ? Depending on what type of account it is it might be a trashcan for deleting an email, or if it's a Gmail account you may currently have it set to as an archive icon : http://support.apple.com/kb/HT4207
What is archiving : http://mail.google.com/support/bin/answer.py?answer=6576

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