550 Requested action not taken: mailbox unavailable (from website form)

Hi there. I hope somebody can help me with this one.
We are running an Exchange 2010 server. We also have a website hosted externally where users can submit info via a form. The website form sends an email to [email protected]. It has been working fine, but all of the sudden, emails from the
website are not coming through. Emails sent to [email protected] from anything but the website work just fine. The error message is "550 Requested action not taken: mailbox unavailable."
I am at a loss as to how to fix this. This happened a couple of months ago. I spent hours trying to fix it, when *poof*, it just seemed to fix itself. I couldn't tell that anything I had done even fixed it(and I hadn't done much).
Here is the full text of the email error message:
From: Mail Delivery System [[email protected]]
Sent: Friday, December 27, 2013 11:19 AM
To: INFO
Subject: Mail delivery failed: returning message to sender
This message was created automatically by mail delivery software.
A message that you sent could not be delivered to one or more of its recipients. The following addresses failed:
<[email protected]>
SMTP error from remote server after RCPT command:
host mail.domain.com[76.164.20.91]:
550 Requested action not taken: mailbox unavailable
--- The header of the original message is following. ---
Received: from icpu2798.perfora.net (infong530.lxa.perfora.net [74.208.16.79])
by mrelay.perfora.net (node=mrus0) with ESMTP (Nemesis)
id 0Mb6xv-1WCEwr3vhL-00KIuD; Fri, 27 Dec 2013 14:19:15 -0500
Received: from 64.122.5.254 (IP may be forged by CGI script)
by icpu2798.perfora.net with HTTP
id 3ovjx0-1VVVbA3OuE-011ONA; Fri, 27 Dec 2013 14:19:14 -0500
X-Sender-Info: <[email protected]>
Precedence: bulk
To: [email protected]
Subject: Update Owner Info Request
Date: Fri, 27 Dec 2013 14:19:14 -0500
From: Update Owner Info Form <[email protected]>
Message-ID: <[email protected]>
X-Priority: 3
X-Mailer: PHPMailer [version 1.73]
MIME-Version: 1.0
Content-Type: multipart/alternative;
boundary="b1_8695b5d02b44c4eae95033f81b999a98"
X-Provags-ID: V02:K0:53ln0xw/tvQG8M7ifWOUO4iaRNVVQ/2jTej143S0V/6
GRlTqQmDPYAeqkJzzZiA4c2yn28akfnnD3XGDQuBU4mWjgGGr1
JbNurwD0ZL+1a8T6O0hfhVwyZxiaGZkIvYdFjRMVb0p/mZ+WTl
TAIXGQHL7ov8pwmIfVwRiNuxFnOWignDb8VLJNIJ43klDr0wwE
BDC/pJUTxAPor11ikmcqgIvBhdnHVYNHCcigEDuIl4hKcFQ+B8
cxowzMn5zTNz5Esbu7GGlkk6icWV6SFRCSiwqMTtqV1YoawRhW
ikXUAyVFnBfZLScBES/PmoDItpJxmdE2/+MDZVzSZaZAApTRss
fNDXreUJEoLfl5YttLeUY1PSkaLg1+6m1mX2RwGZFnVpTL34z5
p4C4oknHf0lNmoaYTD/H7848a4Bu67bNVE=

Hello,
Thank you for your update.
If you have any othere question, please free let me know.
Cara Chen
TechNet Community Support

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550 Requested action not taken : mailbox unavailable

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550 Requested action not taken: mailbox unavailable (from website form)

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