After Effects wont let me scrub through audio or render it in one project, why is this?

Only 1 project has this problem when I open it up. It uses more content than others, but when I delete that content and leave just the audio in, it still wont play. Then I open a new project and insert a music track... and suddenly there's sound. Obviously I'm using ram preview with sound, and output options are correct, and it's not muted... I have 16gb of ram on my macbook pro, so having too many resources can't be the problem.

It's mp3, but when I created a new project and imported this audio file it was fine, just not in my original project.
Forgot to mention that when I open the project it says there are 3 missing effects, I started the prject on my PC and brought it over. But when I did that, the sound still worked... while some plugins obviously didn't carry over. Says it's missing a font family from text layer too, but that didn't effect anything last time either. Iswitched the fonts just in case it was and deleted those effects, but nothing changed. Whatever happened the last time I opened this project months ago, something happened that caused this.

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