AirPort does not detect networks, unless another network connection is made

I've just done a clean install of OS 10.5 to see if this problem was software related, but it persists.
AirPort does not detect any network the computer is near (tested in various locations) normally, but when connecting to a network via Ethernet or to another Mac via Firewire to use network sharing, it can update and load web pages as normal. A strange quirk I've noticed though, whenever connecting with ethernet or firewire, AirPort will suddenly detect the network! It still can't quite connect to WiFi properly though. While still connected through firewire or ethernet, it will have no connectivity through a self-assigned IP. If it is disconnected from ethernet and firewire again, the wireless network will still show up in a re-scan for networks but it will fail to connect.
I've opened the computer up to ensure that the airport extreme card is seated properly as well, and that the antenna cable is secured properly.
Any ideas as to what the issue is? Should I look into replacing the AirPort Extreme card?

Got no answer on my original post, so I went ahead with ordering a replacement AirPort card on eBay (item #130487488014). Once installed, the iMac was able to detect and connect to networks normally... but was still unable to load any web pages. I tried on secured and unsecured networks as I did previously, and plugging directly into the router and into my Macbook Pro via firewire, with the same results as before. Safari loads pages when connected with an ethernet or firewire cable, but will not load pages on WiFi alone. I can attempt to access a router setup page via WiFi, and am prompted for a username/password, but the page itself never manages to load.
Any clues as to what may be going on here?!

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