Alert sounds no longer work in 10.9.1

I'm not sure when it started happening, but I can no longer hear alerts or system sounds through the speakers of my MBP.
Other audio, like music, videos, sounds on web pages etc. all work fine.
However, if I do something that would invoke an alert beep, the screen flashes like it's on mute.  I have gone to sound control panel, tried several alert sounds, adjusted the volume, tried different sound output devices, etc.
I've never come across this.  Any ideas?

I have this same question.  The same thing is happening to me on Mountain Lion (10.8.5) on my MacBook.  I get all of the sounds in Mail, but none of the alert sounds anymore.  I tried shutting my computer down and restarting, but it didn't help.  I can't even get the alert sounds to work on the System Preferences page (like when you click on the sounds to sample them).  This never happened with Snow Leopard!

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