All Linksys equipment stopped working! But have internet with NON LinkSYS equipment

My setup is eccentric but has worked for very long. It stopped working Friday. Let me first say there is a reason why I set it up this way. The reason for my setup is based on location of various IP equipment and when I originally purchased the equipment (in case you are wondering why I didn't go a different route). All routers and network card/devices are Linksys. The only non LinkSYS device is a Netgear Print server and 1 PCI card installed on a 2nd PC.
Here is my setup: From a high speed modem, into a wireless BEFW11S4 (4 port router). Port 1 unused. Port 2 to a second wired router, (for DIRECTV VOD and future VUDU) on the other side of house. Port 3 to a local PC. Port 4 to wireless router WRT54G on the far end of house (I used one of the ports not the WAN connection). Wireless is active for both routers. DHCP is disabled except for main basement router. I treated all routers beyond basement one as gateways per say.
My problem: I have several PC's with wireless cards/ devices. Only one working is 1 XP laptop Dell, as noted below (non Linksys) and does so wirelessly and also with a wired connection. The Directv also works from wired router (2nd in line). The Wii also works properly from either wireless router. Of course my PC to the first router also sees the internet. All locations see both routers with 80+% signal. My house is small, but do to wiring limitations, this is the best solution to get internet.
MY PC setups, models given are Linksys except as stated:
1 XP PC w/WUSB54G (USB card)
1 2000 Pro w/ Netgear WG311 (PCI card) Very old PC and does not see internet, it does search and find both routers with high signal. Internet has worked in the past, although not used for some time. I can possibly assume it just does not work or may also be related to this issue.
1 XP Dell with internal wireless (working fine!) Will work with either wireless router and also communicates with print server successfully.
1 2000 Pro w/WPC11 PCMCIA card
1 XP with Linksys PCI card (don't have model number handy)
I also changed ports of the wired equipment to verify it wasn't a bad port. I have internet connectivity on non Linksys equipment. Some of the computers I use daily, the others have not been turned on in a very long time. After problem arose, I decided to check these computers too. I have not made any changes to the setup. As stated I came home and noticed no internet and went to troubleshoot issue. I have used the Linksys software on all products listed to setup the network.(Original disks that came with each equipment) It will show it can see the routers, but shows NO internet connectivity, but all show it sees either router, but does not see the internet. I must stress that I have not altered any of the setup, it just stopped without warning.
At first I assumed it may just be the wire to one router. (The WRT45G). I have also unplugged all equipment, renewed IP, restarted PC, and also repowered modem. What is bothering me is that the DELL is working fine, including being able to print, the Wii is fine and talks out, and I can renew IP and download VOD from the second wired router and DIRECTV reciever. I also assumed that maybe it was an issue with a new Microsoft patch but a few of the PC's have not been on in months, well before this problem arose. Thank you for taking the time to read this lengthy post. I wanted to be as complete as possible. Any suggestions or ideas is appreciated. I have not found any solution on this forum or off of the internet.
NOTE: I also reinstalled software to 2 PC's including reentering my WEP. This has not solved issue either. Problem remains the same. I have also checked internet properties and network configs (on those PCs) although I cannot imagine every PC has changed itself.
Solved!
Go to Solution.

It does not seem to be an adapter or any of your computer's issue, from what you have described I would assume that your router is experiencing some connectivity issues...You should reset both your router's and then re-configure them both...
As you explained that one computer to the first router connects to internet, you may download the first router's firmware and upgrade it...You may download the firmware from here
Follow these steps to upgrade the firmware on the device : -
Open an Internet Explorer browser page on a computer hard wired to the router...
In the address bar type - 192.168.1.1...Leave the username blank & in password use admin in lower case...
Click on the 'Administration' tab- Then click on the 'Firmware Upgrade' sub tab- Here click on 'Browse' and browse the .bin firmware file and click on "Upgrade"...
Wait for few seconds until it shows that "Upgrade is successful" After the firmware upgrade, click on "Reboot" and you will be returned back to the same page OR it will say "Page cannot be displayed".
Now reset your router :
Press and hold the reset button for 30 seconds...Release the reset button...Unplug the power cable from your router, wait for 30 seconds and re-connect the power cable...Now re-configure your router...
Similarly upgrade your second router and upgrade it's firmware...I think after this you should be able to get connection on all the computers...

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