All of today's emails disappeared when I sent a response

This happens to me occasionally. When I'm responding to an email I received, once I click send, I got an error message and the entire list of today's emails, including the one I responded to, disappeared. I rebooted and still no emails from today are listed. I don't even know if the one I responded to got sent. Where did they go and how do I get them back? And finally, how do I correct this???

Not certain, but this can fix myriad Mail problems...
Safe Boot from the HD, (holding Shift key down at bootup), it will try to repair your Disk Directory while the spinning radian is happening, so let it go, run Disk Utility in Applications>Utilities, then highlight your drive, click on Repair Permissions, then move these folder & file to the Desktop.
Move this Folder to the Desktop...
/Users/YourUserName/Library/Caches/Mail/
Move this file to the Desktop...
/Users/YourUserName/Library/Mail/Envelope Index
Reboot.
If that doesn't help & you can afford to redo any Mail Rules...
Trash these files...
/Users/YourUserName/Library/Mail/... MessageRules.plist and MessageRules.plist
/Users/YourUserName/Library/Mail/... MessageRules.plist and MessageRules.plist.backup

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