Apple iMac (colorfull old ones) w. OS X 10.4.11 and OS X 10.7.2 Lion Server

Dear all,
our Apple Network consists of one Apple Lion 10.7.2 Server based on an Xserve Xeon Server, some very New iMacs running Lion, some eMacs running MacOS 10.5.8 and 10.4.11 and some old iMacs running 10.4.11. We're using OpenDirectory, Filesharing, AFP, DHCP, DNS as Network Services.
Now since we've upgraded from Leopard Server to Lion server we've got the Problem that our old iMacs running 10.4.11 somehow aren't abled to authenticated against OpenDirectory any more... So if a user tries to Login at the Login screen against the server and he can't login (at the old iMac running 10.4.11). At the eMac side also 10.4.11 everything works fine. Regarding the system.log of the old iMac it says at some point "Input / Output Error".
Even a fresh 10.4.11 Installation on the old iMac didn't solve the problem.
Any suggestions what to do and how to fix that problem ?
Thanks,
Markus

First, see if your Mac is compatible with Mountain Lion > http://www.apple.com/osx/specs
If it's compatible, first you have to upgrade to Snow Leopard, so buy Snow Leopard > http://store.apple.com/us/product/MC573/mac-os-x-106-snow-leopard That's because OS X Mountain Lion requires Mac OS X Snow Leopard or Lion.
Then, make a backup of your files, insert the Snow Leopard DVD and upgrade to Snow Leopard. After upgrading, go to  > Software Update to update to Mac OS X 10.6.8.
This update will install the App Store on your Mac, so open it and purchase OS X Mountain Lion. While Mountain Lion is downloading, check if your applications are supported > http://www.roaringapps.com
When the download finishes, the Mountain Lion installer will show up, so follow the steps to upgrade OS X

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