BIG Question Mark in Finder Windows

My Hard drive Finder Windows (any programme) OS 10.4.6.
are suddenly showing a large Question Mark between the 'view buttons' and the 'action wheel icon'.
When 'clicked' the question mark gives a message "The Item Cannot be Found".
All items in each individual folder within the window open normally.
Why is this suddenly happening, and can I get rid of the Question Mark somehow.?
Many thanks.
Lance. UK
iMAC FP. Mac OS X (10.4.6)

Before there was a "sidebar", OS X Finder had a "toolbar" where a user could drag arbitrary items (files, folders, applications) for quick one-click access. Although it may not be obvious with the brushed metal finish, "Finder" windows still retain this function.
However, if the original item to which the toolbar shortcut points is deleted or otherwise becomes inaccessible, its icon will be replaced with a question mark (the same thing happens with "Dock" icons).
The shortcuts can be removed from the window by holding down the "⌘" key and dragging the icons off the toolbar.

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