Bonjour - Mac A can see Mac B but not vice versa

Hi
Very odd behaviour with Bonjour. Up until yesterday, my macbook pro (call it A) would see my mac pro (B) - it would appear automatically in A's sidebar, I could listen to B's iTunes library on A and so on. No problem and there hasn't been for a couple of years. B could not see A - but I didn't really need it to and this wasn't an issue.
Suddenly, this morning, the situation is completely reversed. A can no longer see B at all, but B can see A.
I can connect both ways by using 'Connect', I can ping both ways, but I can't get A to discover B using Bonjour.
The only think I've done that I can think can have caused the changes is to use Omnifocus sync (with B holding the sync server) - it worked fine all day, but overnight Bonjour seems to have reversed itself as described above.
Bonjour Browser shows me that B can see A, but A can't see B. I have identical settings on both machines for sharing, network, firewall (all everything).
Bizarrely, for one two minute period a couple of hours ago, the situation reversed itself (A could see B but not vice versa), but then immediately went back to B can see A, but not vice versa.
Both computers are running the same version of Leopard (10.5.7). I've reinstalled the OS on B, just to see, but this hasn't improved matters.
I can find nothing on google or this discussions forum to explain this bizarre behaviour - most support threads seem to peter out without being resolved. Has anyone any idea what's going on please? I'm quite happy getting stuck into the command line, if that's needed.
Many thanks
David

The Macbook Pro is connected wirelessly to the router, the Mac Pro connects to the same router by ethernet.
As far as I'm aware, I'd made no changes to any of the setups (computers or router) before the problem occurred: bonjour worked fine, but in one direction only (the MBP could see the Mac Pro, but not vice versa). Now, the Mac Pro can see the MBP, but not vice versa - and this is a problem because my printer and itunes server both sit on the Mac Pro.
I've set all the firewall and security settings I could find to transparent access - and (except for the obvious difference between the wired and wireless links), the setups are identical on both computers.
By the way, I can connect to the Mac Pro easily enough and transfer files, I just can't see it via bonjour or use Screen Sharing (again the settings haven't changed since they were working!)
Many thanks
David
Many thanks
David

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