Browser Audio a Mystery

We're using Quicktime Pro 7.5.5 in Mac OS 10.4.11 on a G4 MDD 1.42 DP PPC.
The audio-out unit is a USB DAC that is set as the "default" and "output" unit in both the systems audio preferences and the audiio MIDI utility.
Everything is sending to the USB DAC unit, system alerts, .aif and .dv files on the desktop, everything except any of the internet browsers - Safari, Firefox, Opera.
The audio from any of the net-browsers is still being sent to the G4 audio card.
What is wrong here?
Why is Quicktime sending browser audio to the G4 sound card when all settings and defaults have the USB DAC showing as selected?
Thank you.

( update )
The issue is -
incoming mail alerts, mouse-click miscues, .dv files (Quicktime Player) on the desktop, .aiff files off firewire drives in their folders, Audio/Video applications -
all audio is successfully being fed to the USB DAC stereo unit,
EXCEPT the audio from any browser (Safari, Firefox, Opera), which is somehow "successfully" still routing to the tower's sound card.
All possible preferences for audio output selections show the USB DAC (System Audio / Audio MIDI).
Apple Technical says they "have no knowledge" and that the browser audio should be routing to the default audio device.
We've re-installed, trashed preferences, run OnyX, repaired permissions and are fast coming to the conclusion that this non-hardware issue is exactly as it appears, a OS 10.4.11 browser audio issue.

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