Can 1st and 3rd slots of X99S SLI Plus be used for SLI?

Can I use the 1st and 3rd full size PCI slots for SLI on the MSI Global X99S SLI PLUS mobo?
Specs:
* i7-5960X, intend to OC
* Corsair H100i cooling
* Corsair AX860i pu
* 32gig 2400 DDR4
* Samsung XP941 256GB Ultra M.2 PCIe
* 2x Gigabyte GTX 970 Gaming, intend to OC
I'd like to not put the 970's on top of each other when there's so much extra space - will i get PCIx16 if I use the 1st and 3rd full size slots? Any downside at all?

yeah its a shame, makes for a bad design if using sli on air imho, one card runs 20-25c hotter than the other :(
if I had known of this design problem i would have gone for a different board :(
can't understand MSI's reason for this?
not looking forward to the summer heat coming, might have to try water cooling (if there is room) but even that looks a tight fit?
has anyone run wwater cooling on the MSI GTX970 Gaming on these boards?

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