Can i plug an usb drive into a iMac and access it with a Macbook pro?

I have an iMac that is less than 3 months old and my wife now has a two week old Macbook Pro.
IMac:
21.5" screen
3.1 quad core i7
16 GB Ram
1 TB Fusion drive
Macbook Pro:
13" screen
3.0 dual core i7
8 GB ram
512 GB Flash storage
Both have Mountain Lion.
Both are running iOS 6.1.2
I would like to attach an external USB hard drive to the iMac and allow the Macbook Pro to be able to have full access to just the external USB hard drive.  Either it can't be done or I am just not getting the Sharing settings set correctly.  I played around with sharing last night on both machines.  When I would plug in a FAT 32 thumb drive into either machine and find it, I would add it to he shared folder but it would only allow me and "Everyone" to have access.  I could not change "Everyone" to "No Access" because "Everyone" was greyed out or add my wife.  I can access the Macbook Pro as Mark with the iMac but I can only access the iMac from the Macbook Pro as a "Guest".
I am using a Netgear WNR3500L router.  I currently have the USB hard drive connected to the router via the Readyshare USB port but it is formatted as NTFS for Windows shared files.  This router does not recognize HFS+ formatted hard drives in the USB port.   When I move it to the iMac, I will back it up and then reformat it as HFS+.  Since the router is about a year old and working fine, I would buy an Airport Extreme or Time Capsule if that is my only choice.
I don't normally turn off the iMac.
I would not want to allow access to any other device.
Since I am very new at Mac's, I would appreciate any help that any of you can offer me.

Topher Kessler,
Thanks for commenting to me.
My thumb drive is formatted as Fat 32 so your first paragraph explains what I am seeing.
You are correct that my existing router does not support HFS+.  It is visible from my MAC and some file are read only.
The drive that I will be wanting to attach to the iMac would be formatted as HFS+ so it would not be accessible from my router.
So, if I setup sharing (If it is possible) and thern plugged in a USB 3.0 hard drive into my iMac, would it be accessible to my Macbook Pro?  If it would not be, it looks like an Airport Extreme or a Time Capsule are my only options?
I see that a Netgear 4500 is Time Machine compatible.  Any idea what that means?  When I read pages 80 through 82, it appears to me that you format your drive to HFS+ and then eventually plug it into the router usb.
http://www.downloads.netgear.com/files/GDC/WNDR4500/WNDR4500_UM_08Jul12.pdf
Thanks for your comments.

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