Can no longer reply to any emails !?

I am having a problem with Mail in that I can no longer reply to any email. Whenever I click reply I just get the spinning beach ball and then have to force quit Mail. This happens on any email I recieve ??
I can still download and send new emails okay, this only happens when I click reply to respond to an email.
I have repaired permissions and verified my disk several times, and each time no erros are reported ??
The only things that have changed recently is that I have installed an Airport Extremem replacing my old Netgear wireless router, and have downloaded the latest security patch and 10.4.10 update ??
Any ideas would be greatly received.
Thanks
-Mic

What type of mail account is this (POP, IMAP, .Mac)? If IMAP or .Mac, what are your Preferences > Accounts > Mailbox Behaviors > Drafts settings?
Go to Apple Menu > System Preferences > Network, choose Network Port Configurations from the Show popup menu, and make sure that the configuration used to connect to Internet appears at the top of the list. Leave checked (enabled) only the port configuration needed to connect to Internet and Built-in Ethernet (in that order if not the same), uncheck (disable) the rest of network port configurations (Parallels in particular, if you have that installed) and see whether that helps — if it doesn’t, turn ON again the ones you want enabled.

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