Can't send emails after Mountain Lion update

After I updated from Lion to Mountain Lion, I could receive messages from my service provider's account, but I could not send messages. Mail Connection Doctor showed the information "Could not connect to SMTP server" for both SMTP servers, and when I try to send a message, I see that each SMTP server now has "(Offline)" after its name. I can still send from my iPad to the same servers, and also from the service provider's webmail. I have rebooted everything (Mac, wi-fi, router) to no avail, and I have followed all advice in Support articles such as TS3276. I have changed absolutely no settings anywhere from the Lion system, and a check of all settings shows that they match the printout I made of the Lion settings. I do not use iCloud for mail, so that server is disabled in Email Accounts, although the system opens iCloud on port 993 for Contacts, etc.
Below is part of the output from the Doctor, which shows an error "503 wrong state for EHLO command" on ports 25 and 587. Researching this message didn't really help me, and, as I say above, I have changed no settings between OS versions, so I assume that something must have changed with Mountain Lion. Any help would be greatly appreciated.

I'm getting the same behaviour with my ISP's mail server:
INITIATING CONNECTION Nov 25 13:26:06.388 -- host:smtp1.bethere.co.uk -- port:25 -- socket:0x0 -- thread:0x7feac3893370
CONNECTED Nov 25 13:26:06.577 [kCFStreamSocketSecurityLevelNone] -- host:smtp1.bethere.co.uk -- port:25 -- socket:0x7feac3c23ae0 -- thread:0x7feac3893370
READ Nov 25 13:26:06.799 [kCFStreamSocketSecurityLevelNone] -- host:smtp1.bethere.co.uk -- port:25 -- socket:0x7feac3c23ae0 -- thread:0x7feac3893370
220 april.london.02.net ESMTP Service ready
WROTE Nov 25 13:26:07.031 [kCFStreamSocketSecurityLevelNone] -- host:smtp1.bethere.co.uk -- port:25 -- socket:0x7feac3c23ae0 -- thread:0x7feac3893370
EHLO pro.config
READ Nov 25 13:26:07.268 [kCFStreamSocketSecurityLevelNone] -- host:smtp1.bethere.co.uk -- port:25 -- socket:0x7feac3c23ae0 -- thread:0x7feac3893370
250-april.london.02.net
250-DSN
250-8BITMIME
250-PIPELINING
250-HELP
250-AUTH DIGEST-MD5 PLAIN
250-DELIVERBY 300
250 SIZE
WROTE Nov 25 13:26:07.507 [kCFStreamSocketSecurityLevelNone] -- host:smtp1.bethere.co.uk -- port:25 -- socket:0x7feac3c23ae0 -- thread:0x7feac3893370
EHLO pro.config
READ Nov 25 13:26:07.735 [kCFStreamSocketSecurityLevelNone] -- host:smtp1.bethere.co.uk -- port:25 -- socket:0x7feac3c23ae0 -- thread:0x7feac3893370
503 wrong state for EHLO command
, investigating...

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Can't send emails after Mountain Lion update

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