Cannot connect iMac to MacBook using Airport

I have a MacBook and an iMac. Until recently I have been able to share files both ways. Now I can see my MacBook in finder on my iMac, but if I try to connect, I get a message saying "failed to connect". I can access the iMac from the MacBook OK. Hope there is someone out there who can help me.

OK.... the is really baffling me...
Here's my setup:
Airport Extreme - constant PPPoe connection to ISP - running 80211n only
iMac conneting to Airport Extreme via ethernet
Macbook connecting to Airport Extreme via 80211n
Airport Express connected to Extreme via ethernet
Macbook Pro and Powerbook and 2nd Express (relay) connecting to Express via 80211g
iMac also connects to Airport Express via 80211g (in addition to direct ethernet connection)
So.... here's the problems....
Macbook and iMac cannot speak to each other. Macbook can see iMac's itunes but gets an error when tyring to connect (unspecified error - make sure port 3689 is open)
Cannot do a 'connect to server' either way between macbook and iMac.
iMac can see and connect to both Powerbook and Macbook Pro. It can also see itunes library on both of thses and vice versa.
HELP!!!

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