Changes to applications and gaming have no effect on WRT54Gv2.2 and restarting breaks routing

If I power off my router and power it back on, nothing on my network can get out and none of the port forwarding works. 
If I log into the router and use diagnostics to ping stuff outside my network it works just fine, but nothing on my network works any longer. 
Also, even when it is working, making changes to the applications and gaming page has no effect.  Old entries on that page still forward to where they did before I made the changes, even though the router will show them going to the new place I told it to go to.  This is extremely frustrating because I eventually try restarting my router which gives the problem I mentioned above that I can no longer use the internet at all from any machine on my network even though my router is connected properly to the internet.
The way I fix this is by loading an old configuration file and then loading back one with my current settings.  This also doesn't always work and it takes a lot of trial and error to get it back to normal.  This has been happening ever since I last upgraded the firmware, which I had to upgrade because automatically updating DynDNS was no longer working.  I can't find the old firmware anywhere on linksys' site so that I could test if it's a bug in the firmware.  This is my last resort before I buy a new router as it's driving me insane.

Thank you ricewind, I installed a slightly earlier verson of the firmware and it appears to be working properly for the moment.  If the problem doesn't come back I'll report back that it's probably a bug in the version of the firmware I was using (which was WRT54GV4.0_4.21.1_US_code.bin)
Thanks again

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