CS4 Illustrator suddenly stopped saving my guides when I save the file? How to fix?

?

Thanks- I find this website rather unwieldy-I will bookmark your link.  I've used Illustrator since 1994 so i don't usually have a lot of questions. But in CS4 I find that all kinds of things suddenly happen when I roll my mouse over the wrong area .
I had put all my guides on the bottom layer and the whole layer totally disappeared when I reopened the file. It happened twice so I finally just redid and locked the templete layer without converting it to a guide.

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