Do I have to make sure the server stays on all the time?

Is it ok if the server goes to sleep?  If it is needed will it wake up?  Will mail or calendar updates be slower if it is?

Is it ok if the server goes to sleep?
Not if you expect it to work.
If it is needed will it wake up?
If you have a Bonjour sleep proxy on the network (such as an AirPort base station), it may wake for access to Bonjour-advertised services if so configured in the Energy Saver preference pane.
Will mail or calendar updates be slower if it is?
It won't wake for those services. If you want to run a server, don't put it to sleep.

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