Does the computers DAC or the iPods DAC work when playing a song on iTunes?

In other words:
Which devices DAC is being used when playing a song on iTunes?
In other words:
When I connect an external DAC to my PC and play a song from my iPod on iTunes, which devices DAC is being used?

iTunes does not "play a song from your iPod", only from the content of the iTunes library.  Therefore, anything played in iTunes will make use of whatever audio processing components are in or attached to your PC.  The iPod's DAC is used only for audio output by the iPod via its headphone jack (or, in some configurations, via the USB cable - e.g., for some in-car setups).

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