Email - How Does It Send Mail Differently compared to a Palm?

I'm curious... and hoping someone can clarify this for me. 
On the Blackberrys we have for our company it does not allow you to specify an outgoing server, but automatically configures one presumedly hosted by our service provider, Bell (and doesn't let you see which one it is using).  Our blackberrys are setup to use work emails (ie. incoming server is our company mail server).   Our spam filter is set to NOT allow people to send from external IP addresses using their work emails, yet the Blackberrys are able to send even though they are using the Bell network/server.  However, on our Palm devices (Palm Preo), we cannot send from a work email address through an external IP, even though on first glance the different devices appear to have a similar setup, except that we can actually see the outgoing server (again, a Bell server, mail.1xbell.ca)  on the Palm.  
Basically, I'm trying to figure out how the Blackberrys handle outgoing mail differently to allow them to bypass the spoofing rule.

Hi and Welcome to the Forums!
You have touched upon one of the fundamental differences between the compared devices. This is how I think it works...
Others: These run an email client on the HH that touches the email server for both incoming and outgoing email, using that infrastructure for traffic both ways. This generates traffic over the carrier network to the HH even when there is nothing to do.
Non-BES BBs: These use Blackberry Internet Service, a service, hosted at the carrier, which functions as the conduit between the internet facing email service and the BB. For incoming, BIS touches the email server. For outgoing, BIS uses it's own SMTP host (at the carrier). Since BIS touches the server to check for new email, no traffic to the HH is generated unless there is something to do.
BES-based BBs: These use the corporate BES for handling email in both directions.
Hope that helps!
Occam's Razor nearly always applies when troubleshooting technology issues!
If anyone has been helpful to you, please show your appreciation by clicking the button inside of their post. Please click here and read, along with the threads to which it links, for helpful information to guide you as you proceed. I always recommend that you treat your BlackBerry like any other computing device, including using a regular backup schedule...click here for an article with instructions.
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