Ethernet not connected when using external hard drive

My brand new Mac Mini (mid 2010) works fine when I connected it out of the box to my router via Ethernet cable. But when I boot that new mini using an external hard drive (La Cie 500 GByte Firewire 800 connection), the Ethernet shows up as "Not connected". It advises that I should plug it in, or the other end may not be plugged in. But when I simply reboot again to the internal hard drive, without changing any connections, the internet connection works fine. When I boot a different and older Mac Mini (2.0 GHz Core Duo) off that same external hard drive (using Firewire 400 of course), the internet connection works fine. I'm stumped despite using Macs since 1986. I tried automatic mode, but I can't get far, because it thinks there is nothing plugged into the Ethernet port. But there is, and it even works when booting off the internal hard drive. Both the internal and external OS are at 10.6.4, latest. I also tried erasing the internal hard drive so it couldn't get confused, but that didn't help. What else to try?

The ethernet service has nothing to do with the cable plugged or not, and the ethernet cable is working fine with the other system, so definitely the issue with your other OS. Forget about the working OS for now and boot into the OS that has the problem and select the ethernet service from the service list in System Preferences > Network then click on minus button then apply the changes then click the plus to add it again.
If this didn't work, go to :
/Library/Preferences/SystemConfiguration
and trash the NetworkInterfaces.plist file then restart and see if it solves the issue.

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