Exchange 2010/2013 coexistance mailflow issues: 421 4.4.2 socket error.

So I am in Exchange 2010 SP3 / Exchange 2013 SP1 co-existence. 
I can send from a test 2013 user to external and 2010 users internally on the domain. But I cannot send to the 2013 test user. I get:
451 4.4.0 Primary target IP address responded with: "421 4.4.2 Connection dropped due to SocketError." Attempted to failover to alternate host, but that did not succeed. Either there are no alternate hosts, or delivery failed to all alternate
hosts.
I've been looking at this for example: 
http://support.microsoft.com/kb/979175
But no matter where on the 2013 receive connectors I add Exchange Server Authentication, it still doesnt work.
Theres so much stuff on this error message, but everyhing I find seems to be 2003 / 2010 or other coexistance which is different to my environment. 

Hi guys, thanks for the responses, please keep in mind I am not a specialist Exchange Admin, I'm a IT jack of all trades.
We do not use Windows firewalls on the domain network. Both my 2010 and 2013 setups are in DAGs. Telnet client is not installed on the Exchange 2013 servers, only on the 2010 servers.
How do I "drop an email...through Telnet"? 
Telnet from SiteA Exc2010 to SiteA Exc2013:
220 Exc2013.MyDomain.local Microsoft ESMTP MAIL Service ready at Wed, 4 Jun 201
4 09:42:39 +1000
451 4.7.0 Timeout waiting for client input
Connection to host lost.
Telnet from SiteA Exc2010 to SiteB Exc2013:
Blank window, nothing comes up, no response at all. Doesnt seem to time out either.
Telnet from SiteC Exc2010 to SiteA and SiteB Exchange 2013:
Exactly the same as from SiteA Exc2010.
IPConfig:
Windows IP Configuration
   Host Name . . . . . . . . . . . . : Exc2010
   Primary Dns Suffix  . . . . . . . : MyDomain.local
   Node Type . . . . . . . . . . . . : Hybrid
   IP Routing Enabled. . . . . . . . : No
   WINS Proxy Enabled. . . . . . . . : No
   DNS Suffix Search List. . . . . . : MyDomain.local
Ethernet adapter Exchange MAPI Network:
   Connection-specific DNS Suffix  . :
   Description . . . . . . . . . . . : BASP Virtual Adapter
   Physical Address. . . . . . . . . : 00-26-B9-5E-E7-47
   DHCP Enabled. . . . . . . . . . . : No
   Autoconfiguration Enabled . . . . : Yes
   IPv4 Address. . . . . . . . . . . : 172.16.2.8(Preferred)
   Subnet Mask . . . . . . . . . . . : 255.255.255.0
   IPv4 Address. . . . . . . . . . . : 172.16.2.31(Preferred)
   Subnet Mask . . . . . . . . . . . : 255.255.255.0
   Default Gateway . . . . . . . . . : 172.16.2.2
   DNS Servers . . . . . . . . . . . : 172.16.2.12
                                       172.16.2.1
   NetBIOS over Tcpip. . . . . . . . : Enabled
Ethernet adapter Exchange Receive:
   Connection-specific DNS Suffix  . :
   Description . . . . . . . . . . . : Broadcom BCM5709C NetXtreme II GigE (NDIS
 VBD Client) #50
   Physical Address. . . . . . . . . : 00-10-18-FC-16-76
   DHCP Enabled. . . . . . . . . . . : No
   Autoconfiguration Enabled . . . . : Yes
   IPv4 Address. . . . . . . . . . . : 172.16.2.15(Preferred)
   Subnet Mask . . . . . . . . . . . : 255.255.255.0
   Default Gateway . . . . . . . . . : 172.16.2.2
   DNS Servers . . . . . . . . . . . : 172.16.2.12
                                       172.16.2.1
   NetBIOS over Tcpip. . . . . . . . : Enabled
Ethernet adapter Local Area Connection* 9:
   Connection-specific DNS Suffix  . :
   Description . . . . . . . . . . . : Microsoft Failover Cluster Virtual Adapte
r
   Physical Address. . . . . . . . . : 02-26-B9-5E-E7-46
   DHCP Enabled. . . . . . . . . . . : No
   Autoconfiguration Enabled . . . . : Yes
   Link-local IPv6 Address . . . . . : fe80::3c54:d53e:e2ea:8d9f%19(Preferred)
   IPv4 Address. . . . . . . . . . . : 169.254.1.173(Preferred)
   Subnet Mask . . . . . . . . . . . : 255.255.0.0
   Default Gateway . . . . . . . . . :
   DHCPv6 IAID . . . . . . . . . . . : 604120761
   DHCPv6 Client DUID. . . . . . . . : 00-01-00-01-13-4C-3C-35-00-10-18-6B-C0-36
   DNS Servers . . . . . . . . . . . : fec0:0:0:ffff::1%1
                                       fec0:0:0:ffff::2%1
                                       fec0:0:0:ffff::3%1
   NetBIOS over Tcpip. . . . . . . . : Enabled
Ethernet adapter Exchange DAG Replication:
   Connection-specific DNS Suffix  . :
   Description . . . . . . . . . . . : Broadcom BCM5709C NetXtreme II GigE (NDIS
 VBD Client) #49
   Physical Address. . . . . . . . . : 00-10-18-FC-16-74
   DHCP Enabled. . . . . . . . . . . : No
   Autoconfiguration Enabled . . . . : Yes
   IPv4 Address. . . . . . . . . . . : 10.10.2.8(Preferred)
   Subnet Mask . . . . . . . . . . . : 255.255.255.0
   Default Gateway . . . . . . . . . :
   NetBIOS over Tcpip. . . . . . . . : Enabled
Ethernet adapter Backup Network:
   Connection-specific DNS Suffix  . :
   Description . . . . . . . . . . . : Broadcom BCM5709C NetXtreme II GigE (NDIS
 VBD Client) #5
   Physical Address. . . . . . . . . : 00-10-18-6B-C0-36
   DHCP Enabled. . . . . . . . . . . : No
   Autoconfiguration Enabled . . . . : Yes
   IPv4 Address. . . . . . . . . . . : 192.168.2.8(Preferred)
   Subnet Mask . . . . . . . . . . . : 255.255.255.0
   Default Gateway . . . . . . . . . :
   NetBIOS over Tcpip. . . . . . . . : Enabled
Tunnel adapter isatap.{7282FD1F-E6A4-4BD2-8D40-B2586BF4130D}:
   Media State . . . . . . . . . . . : Media disconnected
   Connection-specific DNS Suffix  . :
   Description . . . . . . . . . . . : Microsoft ISATAP Adapter
   Physical Address. . . . . . . . . : 00-00-00-00-00-00-00-E0
   DHCP Enabled. . . . . . . . . . . : No
   Autoconfiguration Enabled . . . . : Yes
Tunnel adapter isatap.{C38886F3-875D-4403-A95B-C1BF2243D6BE}:
   Media State . . . . . . . . . . . : Media disconnected
   Connection-specific DNS Suffix  . :
   Description . . . . . . . . . . . : Microsoft ISATAP Adapter #2
   Physical Address. . . . . . . . . : 00-00-00-00-00-00-00-E0
   DHCP Enabled. . . . . . . . . . . : No
   Autoconfiguration Enabled . . . . : Yes
Tunnel adapter isatap.{46074087-7F11-4414-8B45-8EE71DA621D4}:
   Media State . . . . . . . . . . . : Media disconnected
   Connection-specific DNS Suffix  . :
   Description . . . . . . . . . . . : Microsoft ISATAP Adapter #3
   Physical Address. . . . . . . . . : 00-00-00-00-00-00-00-E0
   DHCP Enabled. . . . . . . . . . . : No
   Autoconfiguration Enabled . . . . : Yes
Tunnel adapter isatap.{73064E78-05CB-4279-8EA8-3E5094067025}:
   Media State . . . . . . . . . . . : Media disconnected
   Connection-specific DNS Suffix  . :
   Description . . . . . . . . . . . : Microsoft ISATAP Adapter #4
   Physical Address. . . . . . . . . : 00-00-00-00-00-00-00-E0
   DHCP Enabled. . . . . . . . . . . : No
   Autoconfiguration Enabled . . . . : Yes
Tunnel adapter isatap.{4F2BDC5B-35FF-49D7-9431-67FA2EB1D327}:
   Media State . . . . . . . . . . . : Media disconnected
   Connection-specific DNS Suffix  . :
   Description . . . . . . . . . . . : Microsoft ISATAP Adapter #5
   Physical Address. . . . . . . . . : 00-00-00-00-00-00-00-E0
   DHCP Enabled. . . . . . . . . . . : No
   Autoconfiguration Enabled . . . . : Yes
Tunnel adapter Reusable ISATAP Interface {C3216126-6DDC-4523-958A-5907C784EC1F}:
   Media State . . . . . . . . . . . : Media disconnected
   Connection-specific DNS Suffix  . :
   Description . . . . . . . . . . . : Microsoft ISATAP Adapter #6
   Physical Address. . . . . . . . . : 00-00-00-00-00-00-00-E0
   DHCP Enabled. . . . . . . . . . . : No
   Autoconfiguration Enabled . . . . : Yes
Tunnel adapter Teredo Tunneling Pseudo-Interface:
   Media State . . . . . . . . . . . : Media disconnected
   Connection-specific DNS Suffix  . :
   Description . . . . . . . . . . . : Teredo Tunneling Pseudo-Interface
   Physical Address. . . . . . . . . : 00-00-00-00-00-00-00-E0
   DHCP Enabled. . . . . . . . . . . : No
   Autoconfiguration Enabled . . . . : Yes

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    Hi
    One try, please move administrator mailbox to 2013 server
    Cheers
    If you have any feedback on our support, please click
    here
    Zi Feng
    TechNet Community Support

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    If you have any feedback on our support, please click
    here
    Wendy Liu
    TechNet Community Support

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