Exchange - Cannot receive emails but can send

Using Exchange 2007 - (CAS,Hub,Mail and Transport). I have all external mail coming in on http/s to webmail and outlook anywhere and port 25 smtp (mail.domain.com). Firewall NAT rule is External to Internal.
I then created a DMZ and now have webmail and outlook anywhere http/s going from External>>>>DMZ>>>Internal and it works fine. But when I changed smtp 25 to External>>>DMZ>>>Inside - I cannot receive any emails from
external, I can send though.
I Only changed the way it is being routed, instead of going from External (mail.domain.com) to Internal (mail server). I have it going External (mail.domain.com)>>>(DMZ IP) >>>(Internal Mail server).
I didn't change any IPs or URLs or anything on the External or Internal and webmail and outlook anywhere are working fine through the DMZ.
I thought maybe it could be the external mail filter in the cloud that hits our external domain name but since I'm not touching anything external I don't see how that would matter.
Any suggestions? Thanks!

Hi,
Based on my experience, improper permission groups would always bring mail flow issue. Thank you for sharing the solution here.
http://social.technet.microsoft.com/Forums/exchange/en-US/75acd965-3ca5-421f-9994-89657bbddf7d/cant-receive-external-emails?forum=exchangesvrdeploylegacy
And here is a reference about configuring Exchange 2007 Hub Transport role to receive Internet mail:
http://blogs.technet.com/b/exchange/archive/2006/11/17/configuring-exchange-2007-hub-transport-role-to-receive-internet-mail.aspx
If you have any question, please feel free to let me know.
Thanks,
Angela Shi
TechNet Community Support

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