Exchnage 2007 + SP3on Windows 2008 R2 - Cannot Receive Internal e-mail (sent via Outlook or OWA)

I have installed Exchange 2007 + SP3 (All Roles on 1 server) onto a Windows 2008 R2 Server Member Server (after all pre-reqs met)
On Windows 7 client (or Windows XP) I have Outlook 2007 (using cached mode - but also tried with this turned off), and as an AD user with a mailbox created on the Exchange 2007 + SP3 Server I can configure Outlook 2007 to connect to my Exchange 2007
+ SP3 Server and can access the Mailbox (and ran Test on Outlook config to find SCP etc. and this is OK). I can access the GAL and can compose and Send an e-mail. Copy of the e-mail goes into the Sent box (via a Web Browser i.e. OWA mail ends up in Drafts
folder), buy when I logon and access the mailbox of the user the e-mail was sent to, they have not received the e-mail.
Any Ideas ?
Note - With Exchange 2007 + SP2 this unable to receive e-mails is not a problem either in Outlook or OWA

With no messages in the Message Tracker or Queue Viewer I decided to install an additional Exchange 2007 + SP3 Server (all Roles again) onto another Member Server but this time Windows 2008 + SP2 rather than Windows 2008 R2 + SP1
I moved across 2 mailboxes from the original Exchange database to the new Exchange server
I left all other mailboxes hosted on the original Exchange Server
From the same client units I then logged on as these 2 mover users and could now sens and receive e-mail and on the new Excahnge Server could see Messages using Message Tracker.
I could now also send and receive e-mails to and from the mailboxes on the old server to\from mailboxes on the new server.
I powered down the new Excahnge Server and then also found I could send\receive e-mails to\from mailboxes hosted on the original Exchange Server which I could not do previously
Thus I do not understand how the problem appears fixed (for the time being at least) simply by having installed another Exchange Server (only onto a Windows 2008 + SP2 unit rather than a Windows 2008 R2 + SP1 unit)
I will test again tomorrow.
Thanks

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