Extremely slow file server performance

I have a Time Capsule (1TB, late 2008) which I use as a file server. The client of interest is a Mac mini, slightly newer. The connection is 802.11n at 100+ MB/s. I can see no evidence of interfering networks.
Anyway, dragging largeish (350MB) files from the file server to the Mac mini using finder is sometimes incredibly slow (1hour +). Tranfering the same files using firefox, a file:///Volumes... URL and "Save link as" takes perhaps 5 minutes.
General network connection and wider internet access is fine.
Any ideas what the problem is?
Everything is fully updated.

Hi Dennis,
From the description a lot of open files exist which seems to be the cause. Instead of reboot, please test to manually kill these processes in MMC - Share Folders - Open Files and see if the performance will back to normal.
If so, try to find out the exact process which locked these files. Meanwhile please help collect a screenshot of open files for helping us understand the current situation.
Meanwhile we have a forum FAQ which provided most general steps to troubleshoot network performance related issue. Please see if it could help. 
If you have any feedback on our support, please send to [email protected]
I tried to kill all of the open "\msftewds" files. They disappeard and then came back again. I was
unable to kill them.
How can I find out which process is locking an opened file? I'm sorry but at the moment I can't give
you a useful screenshot because at the moment we are fine with the fileserver. As soon as the problem will be back I'll upload a screenshot.

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