Font is italicized, but only in web browsers

Hi everyone,
I have a strange problem.  I've started work with a new inherited workstation, and my font is italicized, but only in web browsers.  And I *think* only on text that has default styling.  E.g., forms, textareas, Google search results, and other unstyled text.
Is there a system-wide setting that changes web browser text styling, or have I inherited a bizarre workstation?

One thing to check if it's only in the Browser, like Safari>Preferences>Advanced, does it have a custom css set there...

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