G4 won't boot after last Security Update

I have an AGP G4 tower that's been running 10.4.11 fine until today. The last thing I did was Software Update. The only things on the list were an iLife update and a Security Update. I don't know the update number but since I always keep the system updated, I can only assume these two updates were the latest available.
So after the update... I hit the "restart" button but then I get the grey screen with a prohibitory sign and a spinning wheel.
It's not a hardware issue... I can reboot this machine into an alternate hard drive just fine.
I fixed permissions.
I ran disk utility - hard disk checked out ok.
Still can't boot using this drive.
Obviously this last security update broke my system folder because it's as if it can't find a valid system.
So now the question is, what can I do to fix my system folder?
Thank-you.

Simple fix.
I downloaded and installed the 10.4.11 Combo Update onto original drive while booted into the alternate drive. I then downloaded and installed the Security Update 2008-0001 onto the original drive. As a precaution I repaired permissions on the original drive.
Successfully booted into original drive.

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