Gutted. Yosemite Mail lost all my signatures.

Gutted. Yosemite mail has lost all my signatures. 100+ time saving replies and standard emails, that have taken years to put together. They were there yesterday, and now only the names remain, the signature contents are blank.
So I used TimeMachine to replace user/library/mail/V2/signatures folder, with Mail closed - but this made no difference. Mail straight away wipes over it. The files remain in the signatures folder.
I originally copied my Mail folder direct from the last system, overwriting user/library/mail. On first launch, Mail took some time to update the folder. So surely this isn't the problem.
Any help gratefully appreciated. Anyone else with the same problem?
Thanks
Jon

Thanks Richard...  I have no idea what the following means!
Last login: Mon Nov 24 11:34:19 on ttys000
Jonathans-MacBook-Pro:~ jon$ syslog -w -k Facility com.apple.mail
Nov 24 11:35:07 Jonathans-MacBook-Pro.local Mail[1570] <Warning>: An instance 0x600000329ce0 of class ConversationMember was deallocated while key value observers were still registered with it. Current observation info: <NSKeyValueObservationInfo 0x618000a46de0> (
  <NSKeyValueObservance 0x6000002c2760: Observer: 0x6100001c4920, Key path: messageDataCached, Options: <New: YES, Old: NO, Prior: NO> Context: 0x1049fb61c, Property: 0x6180018457c0>

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