Has ip address, cannot connect iTunes, Safari

Here is a strange one. One of my 8 macs mostly G4s all running 10.5.8 cannot connect to the internet or iTunes store even though it shows a 192 address shows all the other macs on the network. Software update says everything is current even though the last update it shows installing is April 10th but it doesn't say it cannot connect. So is it really connecting? What could be causing the inability to connect Safari and what would be the cure?
TIA
Enio Cordoba

Thanks Camelot but this is your second suggestion on this topic but both seem foreign to me. (previously you suggested reading logs) Absolutely do not understand  what a web proxy is and there were no other similar posts about proxys to understand this. You previously said to ping the network and read the various logs but this is also new to me sorry.

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