Help -Why did this happen to my messages?

Messages saved in a mailbox were all changed to read:
"The message from [email protected] concerning “Re: Note from Jeri” has not been downloaded from the server. You need to take this account online in order to download it."
How do I retrieve original messages?
Mac G5 2gig dual   Mac OS X (10.3.8)  
IMAC (Intel)   Mac OS X (10.4.7)  

For the Mail.app under which Mac - the PPC running 10.3.8 or the iMac Intel running 10.4.7?
Is this for a user created "On My Mac" location mailbox?

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