How best to Sync 2 Computers to the i-phone?

Can anyone post a step by step guide to syncing two computers to the i-phone. I have a Macbook Pro and an i-mac. I hardly ever use the i-mac, it's mostly my back-up and guest log in computer.
Right now the i-mac has thousands of e-mails that have not been sorted out as I do that mostly on my MacBook Pro. I would like to have both the computers have the same information for contacts, calenders and e-mail, and then have it all sync up with the i-phone, when I get it!!! (Waiting on a DF at&t order!) Thought I might as well get ready.... Any thoughts and suggestions would be much appreciated and of-course helpful to others in the same position....
Message was edited by: seabreeze1982

Sorry neuroanatomist, I edited the question just as you answered.... tried to make it simpler...
The Data on my MacBook Pro is all current, the data on the i-mac is old and the e-mail files are huge on the i-mac as I have never deleted anything on it.... If as they are saying, that mobileme will sync everything, I would like to clean up the -imac and match it to my macbook. Except I recall doing this once and it deleted the e-mails from the server? So I lost e-mails I didn't want to....
I believe that the inbox and the smart mailboxes are e-mails that are on the apple server? If I move them to the "On my Mac" section with folder, they will be saved, if I delete them from the in-box they also get deleted from the server, is this correct?
I just want the sync with the i=phone to be correct, and not put double information on the phone.
Thanks

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