I upgraded and lost a lot of data ...where can I recover it?

I have an imac 27 inch 2009 just upgraded to 10.7.4 and lost mail data and downloads.
Can anyone tell me how to recover my lost files?
Thanks
Dorothy

Do you have a hard drive that's not filevaulted and desperate enough to spend hours and a couple of hundred of dollars in a data recovery effort?
Or could you redownload the files that were in the Downloads folder (and shouldn't have remained in there as it's for temporary downloads only) and perhaps do without your soon to be old emails?
Create a data recovery, undelete boot drive

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