ICloud inbox not syncing to iMac running Yosemite

This problem started when I was running the Yosemite beta (on a separate drive); my iCloud (mac.com address) e-mail would sync fine on the Yosemite beta, but nothing from the inbox would appear on Mavericks unless I was on the computer at the time the mail arrived. Everything showed up fine on my iOS devices, my Macbook Air running Mavericks and the Yosemite beta mail.
Upgraded to Yosemite hoping the problem would be fixed, but it's not. Nothing in my Mail inbox will sync from iCloud down to my iMac. Everything's there that's in folders, everything shows up on all my other devices, and if new mail comes while I'm on my iMac and Mail is open it will show up. But any other mail in my iCloud inbox will NEVER show up on my iMac. I've tried everything, mostly more than once: turning account on and off; deleting account and setting up again; removing certain files from the Library-Mail-V2 folder (though I might have missed the correct ones I guess) but nothing works.
This makes Mail completely useless to me as the iMac is my main computer and I get most of my e-mail to my mac.com account. The only solution I can think of now is to set it up to forward to one of my non-Apple accounts. Any other suggestions appreciated though.

Finally I had the idea to grab a Time Machine backup of my V2 folder, or parts of it, from before I installed the beta. This worked … sort of. I did get all my iCloud mail, but the problem persists. If my iMac is awake with Mail open, new mail from iCloud arrives, if it isn't, it doesn't. And NOTHING I have tried, and I've tried everything I could find and think of, solves this problem.

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