Installed updates--now Airport Card vanished

Greetings,
I have the most recent 21.5" iMac (3.2 GHz) with the most recent OS installed.
Last night, I installed three updates: iTunes 10, Snow Leopard Graphics Update 1.0, and Security Update 2010-005 1.0. I restarted the computer.
This morning, I went to get online and my Airport was gone. It said "Airport Card Not Installed." This is odd because it's worked fine since January.
I went to Networking, and Airport was off. When I clicked "Turn Airport On" nothing happened.
I tried holding shift and restarting, restarting again, and shutting down. It's dead. My Windows laptop gets internet without a problem.
Some forums have recommended holding P and R while restarting. What is that? Other ideas as to what has occurred?
Thanks!

I recently bought and installed iLife '11 and since then (few weeks ago) I have had intermittent issues with the airport card not working and a message saying 'airport not installed' showing. I mention iLife '11 as it is the only significant change to my system since I bought the iMac. Until now I have never experienced any issues with airport cards on this or a long serving previous iMac. The point is nothing I have done (I've hunted out many discussion threads on this and other forums) has made a blind bit of difference (gone through all of the things so far suggested here - nothing worked). The only solution has been to switch off and try again the next day. So far that has worked but I would rather know why this happens and how to stop it then just be at the mercy of a random error / bug. Anyone else had random failures of airport cards? I have grown to expect Apple equipment to be reliable... having re-evaluate that of late!
I have a late '09 iMac with the latest OS updates installed. Linked to a secure wireless network, shared with a Dell PC (also wireless).

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