Is there a difference between "Spotlight" & the "Search" field?

Hi,
In Lion, is there a difference between searching for a file using SPOTLIGHT . . . or the SEARCH field in the upper left of a folder (if you click your Hard drive.)?
For example, I have a MacBook Pro with a 2nd hard drive inside.  If I don't "Spotlight" the 2nd hard drive - will the SEARCH field then still work?  Or if I DO SPOTLIGHT it - is it using the same search mechanism using "Find" and "Spotlight"?
I'm curious how this works?  Thanks very much,
Larry

Using COMMAND-F or Finder's search field provides a window like this:
If you click on the Kind dropdown menu you will see:
You can select different search criteria for locating files. If you then select Other you will see:
You can then check the different search attributes you wish to add to the previously shown dropdown menu thus providing many different search criteria.

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