Itunes 10.5 wont connect to the internet

I recently updated to itunes 10.5. Until then, itunes would connect to the internet ok. now it wont connect at all.
I have read through other forum topics for an answer. I have a firewall but I have checked the firewall settings and itunes is set for full access. i am not using a proxy. I keep getting a message saying my computer is not connected to the internet. This is not correct as all of my other application connect ok to the internet. I have tried completely uninstalling all apple software and then reinstalling but still get the same problem.
How can I resolve this?

Have EXACTLY the same issue thePOPEofDOPE420, tried the setting> general> reset> reset network setting suggestion - but this does nothing. Still the issue persists.
WiFi - internet works great. As soon as it's off, status bars showing a full signal, 3G but the web page/Facebook app/App Store just hangs and won't load.
Not sure who to contact - T-Mobile or Apple?

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