Itunes not recognizing songs in Media Folder

I have my Itunes Media Folder on an external HD.
When I try to play some tracks in Itunes, they have the ( ! ) next to them. A dialogue comes up that says Itunes can't find them.
But I know they're right there on the external drive.
When I choose "locate" I can easily find them.
But Itunes cannot.
Help. Thanks!!!

It's very connected. And Itunes>preferences>advanced shows the drive and media folder.
I can play the tracks from the drive, in the finder.
Drive has same name as always, and folder structure is same as always. thanks. what could it be?

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