Mac Mail & Open Dir Network Accounts

As a system administrator for my family business I have a question.
I am running a Mac Mini server with all services enabled from lion server..
I currently use a Airport Extreme for routing and port fowarding
The client end computers consist of a G4 tower running leopard and a iMac 2009 running snow leopard...
Sometimes when a user connects to his or her network account and tries to send email using mac mail, the messages will either take an extremely long time to send or not send at all. Also sometimes the network accounts begin to act very strange and a reboot is required on the whole server/client side to restore proper function.
Ideas to what is causing this problem?
Joe

Wow, thanks a lot, I think I got it working now.
Is there a website / wiki / knowledge base where I can find all the differences between Snow Leopard Server and Lion Server?

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