Mac Pro Won't Boot, Hard Drives Are Fine, Passes Hardware Test

I have a Mac Pro 1,1 that will not boot anymore.  The hard drives are just fine.  I put them into another Mac Pro 1,1 and they started up fine and that computer is running great.  I have replaced the RAM, put in new Hard Drives, I only have a keyboard, mouse, and display plugged in to it, and it still won't boot.  What happens is this:
Boots up, shows the gray screen.  Shows Apple Logo.  Goes to Black with mouse cursor, stays at the black screen.  I can move the mouse cursor as normal.  Sometimes it turns into a color wheel, but most of the time it's a normal pointer.
I have run the hardware test several times, both quick and extended.  Everything always passes the tests.  I can boot into target disk mode and use the disks fine via firewire.  When I try to boot from a Lion USB disk, or a Snow Leopard DVD, it always does the same as above and stops at the black screen.  Basically, once it gets to the point where it loads an OS X system, it stops at the black screen as if something isn't working right.  Any thoughts?

once it gets to the point where it loads an OS X system, it stops at the black screen as if something isn't working right.
That is the point at which the graphics card starts the Mac OS X loaded graphics Driver, and it may indicate that your graphics card is broken.
Try Safe Mode (hold down Shift at Startup). It takes five minutes to check the Boot drive, then display a login screen. You need your username and password to proceed, even if you normally auto-login.
Safe Mode uses only "simple" graphics. It does not load the graphics driver.
If you need a replacement card, I recommend you skip over all the refurbished and repaired old graphics cards and install nothing older than the Apple-Firmware 5770, about US$250. It works in every model Mac Pro, and drivers are in 10.6.5 and later.

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