Mac won't boot unless I boot in target disc mode firewired to another mac

I've worked for Apple and I am stumped....all of the genius crew at my store are stumped...and the only thing anyone can recommend is to depot my machine! Here is the situation:
The computer froze up, so I restarted- upon waiting for it to restart, after a long while I received the Slashed Circle at boot. When it was looked at in store, we discovered that when the computer is FireWired to another mac, I can boot my computer through another computer with no problem at all (Booting while holding down T on my computer and Booting the secondary computer holding down Option).
They tried to pinpoint the problem, but the computer still acted the same way (not able to boot on its own, but able to boot through another mac) regardless of the RAM or the Harddrive. What do you guys think is wrong with the logic board that would cause this problem?
[2009 15inch Macbook Pro running Mavericks]
Thanks!
Ash

You have me stumped as well.
When you connected your 2009 MBP as the Target,  the display shows the Firewire logo.  Correct?
The other Mac is booted and has access to the 2009 MBP HDD.  At that point, how did you boot the 2009 MBP?    Did the display on the 2009 MBP replace the Firewire logo with a startup display?  Where you then able to operate the 2009 MBP independent of the host Mac?
As I write this response, I have other applications running in the background that prevent me from turning off this MBP and testing Target disk mode with my other MBP.  Let me know how you did it so that I can run a test tomorrow and perhaps find some answers.
Ciao.

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