Mac won't recognise passwords

I can no longer access sites requiring personal passwords (email, facebook, ebanking etc) that I could before. I get to the internet site okay but when I'm prompted for username and password, the computer goes no further. I work through a server network and this has an access password, but it did not hinder other access.
Can't think what could have happened. I did run Tech Tools to clean up directories and optimize HD but have also done that before with no problems.
Have cleared caches in all libraries, and know that this problem occurs with both Safari and Firefox so it's not search engine problems.
Can anyone suggest ideas?

Hi,
I created a new user in Accounts, and rebooted, logged in as new user and tried to access some of the password protected sites. Unfortunately, no joy, which leads me to believe that re-installing the original system from the system disks I have won't solve the problem either.
Have also tried again to verify and repair file permissions using the Disk Utility application but that hasn't worked either.
Looks like Plan E or F by now.
Thanks for staying with this.

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