Machines losing connection to the Network

We are experiencing some irregular network issues whereby we have 2 G4 Mirror Door Desktops acting at servers (10.3.9) and a variety of Powerbook G4s and Macbook Pro's (10.4.x) which are continually dropping off the network sporadically. Apart from these two desktop servers, all the machines affected are laptops strangely enough.
Firewall is not enabled on the Macs.
The Macs lose all network connectivity including printing, internet connection and server connection.
I have tested pinging to and from the Macs and the results are always as though they are off the network. The Macs can however ping itself indicating it can see a network. Whe you remove the machine from the network and re-add it, the IP address range is 169.x.x.x. This is because by default, Apple allocates the user with an IP range so far off the network that its indicating it can see a connection (ie ethernet card and cable) but cannot see the network.
To get the machine back on the network, you need to remove and re-add the network cable to get connection again. We looked into every possibility inclusing Firewall or DNS issues.
We have compared the switches and vLANs and they are all the same; we have checked that Appletalk is broadcasting correctly through the switch; we have added the machines with a static IP address (servers have static IP addresses anyway)and the machines still drop off; we have altered DHCP lease renewal to a longer period and it still dropped off; removed Appletalk from a machine and it still dropped off.

It may be a 'typo' by you, but forget about 'appletalk' - this is not relevant to your problem (and pretty well redundant now). This looks like a TCP/IP issue.
What is doing your DHCP? Presumably you have checked to ensure that there is only the one dhcp server in your network, and another has not accidentally been enabled (on a server, router or airport). What is the dhcp range for the DHCP server?
Your post suggested that the servers, with static IPs, also 'drop off' the network. So a server IP cannot be pinged from a lan machine which has a correct IP assigned to it (rather than one which has a 169 address which obviously will not be able to ping it) ?
On your laptops, when connected via an ethernet cable, have a look at Network Prefs-> Show: Network Port Configurations. In what order are these? Specifically, I am wondering if Airport is getting prioritised above Built-in Ethernet and maybe there is a 'rogue' airport network which the laptops discover and attempt to join ahead of built-in ethernet.
-david

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