Mail 7.3 Accounts Off-line

A domain name recently expired and was renewed after three days. Two Mail accounts related to that server stopped working in Mail, which shows the dreaded 'account off-line' message.
Both incoming and outgoing mail servers are described as not working.
I have manually re-set in-coming and out-going ports and passwords, to no avail. I have taken all accounts off-line, waited a few minutes, then set them all to on-line. I have quit and re-started Mail.
I can access these accounts using web mail from a browser. I have contacted the administrators of the commercial web host and they say these mail accounts are working normally.
Apple Mail seems to be the culprit.
I can delete the accounts and re-make them, but there is a Lot of e-mail in the problematic accounts that would be lost.
Suggestions?

All right: if Mail tells you:
The server returned the error: The server “mail.domainname.com” refused to allow a connection on port 110.
you can verify the truth or otherwise of this statement by opening a command Shell and typing:
telnet domainname.com 110
If the server is responding to connection requests on port 110 you will see something like this in the Shell:
Connected to server.ISPdomainname.com.
Escape character is '^]'.
+OK <[email protected]>
This message confirms that port 110 is working. Unfortunately, it also confirms that Apple Mail is not telling the truth.
My ISP's server is allowing me to connect with it on port 110. Mail is not working, nor is it telling me the truth.

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When you have the browser open (Internet Explorer) then press the Alt key on the keyboard. It should show you the file options at the top..Then uncheck work offline and check..
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Thank you for all the information you provided about what is going on with Mail on all your Apple Devices. I found an article with steps that I think might help.
I recommend reviewing the article, but I found some steps that sound like they apply specifically to what you are experiencing in the section titled "Mail Connection Doctor shows one or more red dots in the account Status column>Click here if your Incoming and Outgoing email account settings are correct, but the issue persists>Click here if a 'Could not connect to this (server type) server…' message appears":
Click here if a "Could not connect to this (server type) server…" message appears
Check for third-party firewall software
Third-party firewall software, such as Little Snitch or Symantec Norton Personal Firewall, could be blocking the necessary ports. If such software is installed, make sure the firewall is not blocking these ports:
ports 25, 465, and 587 (for sending email via SMTP)
ports 143 and 993 (if you use IMAP email)
ports 110 and 995 (if you use POP email)
If you're not sure how to enable the correct ports, try disabling the third-party firewall completely, then test for the Mail issue. Note: The Mac OS X v10.6 firewall does not interfere with ports needed to send email.
Check for Internet Service Provider port blockingYour Internet Service Provider may be blocking the SMTP ports needed to send email.You can use Apple's MobileMe service to find out if your Internet Service Provider is blocking the sending email protocol SMTP:
Open Network Utility in the Utilities folder—in Finder, choose Go > Utilities (or press Shift-Command-U).
Click the "Port Scan" tab.
Enter "smtp.me.com" in the "Enter an Internet or IP address to scan for open ports" field.
Enable "Only test ports between _ and _", and enter 25 for both port fields.
Click "Scan" to test.
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You can find the full article here:
OS X Mail: Troubleshooting sending and receiving email messages
http://support.apple.com/kb/ts3276
Thank you for posting in the Apple Support Communities.
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