Mail and much more lost after software updates

Background - I upgraded from G4 with 10.3.9 to Mac Pro 1.1 with 10.4.7 one week ago.
When I was working on my machine Saturday, I got a software update box. I clicked the button to run the various software updates, and left. When I came to work this morning, I opened Firefox and found all my bookmarks gone. Weird. I opened Mail to check on emails, and it gave me the "this is how you set up your account" box.
Bottom line: I have lost all my email (hundreds of messages, dozens of mailboxes, very bad) and Firefox bookmarks (okay, not so dire, but very annoying).
Question: Why did it do this? What did I do to cause this? Why, Lord, why???
Thanks,
laurieb
Mac Pro 1.1   Mac OS X (10.4.7)  

Maybe I didn't phrase the question clearly. Why did the simple online update (I don't know all what was updated; there's no log to show what was done...) basically zero out/reset all my applications. My fonts are gone as well.
I do still have my working docs as they were on different hard drive from the system and applications. I have email, Firefox bookmarks, fonts, etc. on my old machine, so in a sense, all I lost were 5 days worth of emails and time spent pulling my hair out.
I just want to know what I might have done wrong in the first place to cause this. Just don't want to do it again.
thanks,
laurieb
Mac Pro 1.1 Mac OS X (10.4.7)

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