Mail Problem - can't 'Reply' or write 'New' email message

Hi all:)
I have a bit of a problem with my mail. When I 'reply' to a message, the sender's address does not come up in the 'to' field. I can type in an address and can put in a subject, but the cursor will not go down to the main body of the message. The same thing happens when I try to send a 'New' message - I can input an email address in the 'To' field and type in a Subject, but the cursor will not go down into the main body - consequently, I can't type a message.
Any help would be greatly appreciated.
All the best
PS: I have tried to Rebuild my mailbox but it didn't help with this problem

Hi gav, and a welcome to the forums!
Yes, you can, but Might try this first...
Quit Mail...
First I'd backup these two Mail folders, by right clicking on them in the Finder, then choose Archive/Compress.
Users/YourUserName/Library/Mail
Users/YourUserName/Library/Mail Downloads
Right click on that Mail folder, choose archive, you'll get everything in the folder, and the folder itself in a file called Mail.zip, move it to a safe place, same for the Mail Downloads folder... only the plist is separate.
Try dragging this file to the desktop in case we need to put it back later, then logout/in...
/Users/YourUserName/Library/Preferences/com.apple.mail.plist
(May or may not require you to setup your account(s) again.)
Quit Mail, then In your home folder, try moving this +folder & file+ to the Desktop then reboot...
First, Safe Boot from the HD, (holding Shift key down at bootup), run Disk Utility in Applications>Utilities, then highlight your drive, click on Repair Permissions, then move these +folder & file+ to the Desktop.
Move this Folder to the Desktop...
/Users/YourUserName/Library/Caches/Mail/
Move this file to the Desktop...
/Users/YourUserName/Library/Mail/Envelope Index
Reboot.

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