Mail's Account Directory - accessible from a different start-up volume?

I have two internal hard drives in my PowerMac G4. One volume has Tiger 10.4.11 installed; the other had Leopard 10.5.6. I have not yet fully transitioned from Tiger to Leopard, but I want to do so. However, before I do so I want to know if I will be able to:
(1) run the applications which reside on the Tiger volume when I boot up in Leopard (so that I do not have to waste hard drive space by installing the same applications on two different internal hard drives; and
(2) access the email messages that reside on the Tiger volume when I boot up in Leopard (so that I can keep all my mail in one place, and so that I can access my email no matter whether I boot up in Leopard, or in Tiger, when I need to use older applications that may not be Leopard-compatible and which I do not wish to spend more money on upgrades).
Since this forum addresses Mail and Address Book issues, I would like to get a definitive answer to the second inquiry, namely, if I boot up in Leopard, can I access the mailboxes and email messages that reside on the Tiger volume, or must I absolutely (somehow) bring all those messages into Leopard and be compelled to use only one Mac OS to access the Mail application on its associated start-up drive.
In trying to figure this out myself, I notice that, in the Mail preferences panel, on the Advanced tab of the Accounts button, there is an option called "Account Directory:" that lists "~/Library/Mail" as the path to the directory where email messages are stored. While there is a "Choose" button next to the file path, both the pathname and the button are shadowed out, and I can see no way to activate that option.
I am a little bit confused here, because the appearance of that option implies that one might be able to actually direct the Mail application to store email messages on a volume other than the start-up volume. However, the fact that the "Choose" button and the actual path name are shadowed out appears to cancel out that capability.
Can anyone shed light on this mystery? Is it possible to access email that resides in Library of another volume other than the one used to start-up the Mac, and, if so, how is that accomplished?
bowlerboy_jmb

BowlerBoy_JMB wrote:
I have two internal hard drives in my PowerMac G4. One volume has Tiger 10.4.11 installed; the other had Leopard 10.5.6. I have not yet fully transitioned from Tiger to Leopard, but I want to do so. However, before I do so I want to know if I will be able to:
(1) run the applications which reside on the Tiger volume when I boot up in Leopard (so that I do not have to waste hard drive space by installing the same applications on two different internal hard drives; and
it might work with some self-contained apps that are installed by drag and drop into /Applications but should not be done in general. It will definitely not work with any apps that use installers to install.
(2) access the email messages that reside on the Tiger volume when I boot up in Leopard (so that I can keep all my mail in one place, and so that I can access my email no matter whether I boot up in Leopard, or in Tiger, when I need to use older applications that may not be Leopard-compatible and which I do not wish to spend more money on upgrades).
Since this forum addresses Mail and Address Book issues, I would like to get a definitive answer to the second inquiry, namely, if I boot up in Leopard, can I access the mailboxes and email messages that reside on the Tiger volume, or must I absolutely (somehow) bring all those messages into Leopard and be compelled to use only one Mac OS to access the Mail application on its associated start-up drive.
yes, you must transfer all your Mail to the leopard partition and use it from there. yes, you are compelled to do so and no there is absolutely no way around it.

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